Genuclymenia humboldti (Pusch, 1837) from the P. trachytera  and early L. styriacus Zones at Jabłonna and Kadzielnia in the Holy Cross Mountains.

Genuclymenia humboldti;  growth lines, suture, two stages in development of suture, septal geometry and outline of the shell.

Diagnoza

Umbo covering about one third of the conch diameter; slopes of the flank lobe merging approximately at right angle, the ventral one obliquely crossing the conch flanks.

Porównanie

Probably Ammonites humboldtii Pusch, 1837, based on a pyritized specimen from bituminous calcareous shale exposed at the foot of Kadzielnia hill in Kielce, is the first reference to this species, but Pusch (1837) was uncertain whether the siphuncle is ventral or dorsal. Roemer (1866, p. 675, pl. 13: 1) illustrated a specimen of probable Protornoceras from apparently the same exposure, suggesting that Pusch’s A. humboldti and A. buchi represent variants of the same goniatite species (which seems highly unlikely). Gürich (1896, p. 329) used his collection of specimens, considered by him conspecific and coming probably from the same stratum exposed between Kadzielnia and Psiarnia hills, to prove that this was a clymeniid and transferred the species to Cyrtoclymenia. Although the original Pusch’s collection has not survived to our days, one of the specimens studied by Gürich (1896) was illustrated by Frech (1902, pl. 4: 5). This emendation of the first reviser should thus be binding, so more that it was accepted by Sobolew (1914, p. 64) who distinguished two varieties within Gomi-monomeroclymenia humboldti. The variety G. h. genulobata, based on the material from Sieklucki’s brickpit, with pointed flank lobe of the suture seems to fit within the range of the species, as suggested by additional material assembled by Czarnocki (1989) and myself. However, Czarnocki (1989, p. 64) preferred to abandon Pusch’s species name and to replace it with his Flexiclymenia puschi. He proposed the holotype collected at Ostrówka from the Prolobites Zone strata (thus his beds 1-3) which seems to represent another Sobolew’s species, here referred to as Praeflexiclymenia curvidorsata. I believe that Flexiclymenia tempestiva Czarnocki, 1989, F. staszici Czarnocki, 1989 and F. simosa Czarnocki, 1989 are conspecific with G. humboldti.

Autekologia

Występowanie geograficzne

 Jabłonna; Ostrówka; Kadzielnia and reworked to Quaternary sediments at Sieklucki’s brickpit in Kielce.

Zasięg czasowy

P. trachyter to the early L. styriacus  Zones; probably the L. styriacus Zone; black shale.  

Materiały muzealne

ZPAL: 26 specimens.

Literatura

Dzik, J. 2006. The Famennian "Golden Age" of conodonts and ammonoids in the Polish part of the Variscan sea. Palaeontologia Polonica 63, 1-359.